早期中生代真蕨植物生殖器官构造和原位孢子研究

Through comprehensive investigations, several results and progresses are made in the following three aspects of the project 39900007 "Studies on the Early Mesozoic ferns from China: fine structures and in situ spores": Firstly, the fine structures of the fertile organs and in situ spores of the typical ferns are studied by anatomical and electron microscopical approaches: the in situ spore ultrastructure of the Mesozoic marattialean fern (Marattia asiatica) is investigated using SEM and TEM techniques. Five species of the widespread Osmundaceae fern- Todites are examined and described based on the collections from the Jurassic in Jiangsu and Anhui, including one new species (Todites nanjingensis sp. nov.), the fossil records and geographical distribution of Todites in China are overviewed in some details. The Matoniaceae fern Phlebopteris from the Lower Jurassic in western Hubei is studied and described with emphasis on the fertile organs and in situ spores as well as the fossil distribution of Phlebopteris in the Mesozoic of China. The fertile organs and in situ spores of three Dipteridaceae ferns are reported including Dictyophyllum nilsoni，Clathropteris obovata and Hausmannia sinensis sp. nov. Over 10 species ascribed to 8 genera of associated ferns with the early angiosperms from Late Mesozoic are described. Two new forms of Bryiidae (Musi) are reported from the Jurassic in Juggar basin，Xinjiang. Secondly, as the ecology and climate indicator aspects of the fern are considered, the living habitat and climate tolerance of fern fossils as well as the scientific significance of the ferns being used as the "thermograph of the geological history" are emphasized. A remarkable fern community is recognized from the Early Jurassic in western Hubei, which is a representative floral community in the Southern Floristic Province of China during the Early Mesozoic, indicating a tropical to subtropical climate and a autochthonous peat-forming environment. Preliminary analyses is made on the distribution pattern of about ten climate indicator plants in the Late Mesozoic deposits in China, including a Gleicheniaceae fern Gleichenites. A data base of above plants and its components in the Early Cretaceous of East Asia has been compiled. Thirdly, based upon the update results and our new progress, the dispersed spore assemblages are recognized in the Late Triassic in northern Shaanxi, the Middle Jurassic in Nei Mongol and the Jurassic in Qaidam Basin. According to comparative results with in situ spores, the vegetation aspects are reconstructed and the paleoenvironmental significance is discussed regarding to the formation of coal, petroleum and gas resources. .About 12 articles are get published in the professional journals, including five in international SCI journals and one cooperated monograph, three additional manuscripts are submitted to other international journals. Extensive and positive collaborations are made with universities and institutes of France, Holland, Japan and German. About 11 results were presented and posted in the international and domestic conferences and four other special invited talks were presented in the universities of USA and Germany.

以扫描、透射电镜及超薄切片等方法，深入研究华南西北早期中生代六个科典型真蕨植物的生殖器官构造及原位孢子；查明孢子囊群或聚合囊的微细构造及原位孢子的微观形态和超微结构特征；与化石和现代真蕨植物及分散孢子进行比较；对部分植物体进行复原重建并探讨系统分类和发育演化意义；进一步提高我国早期中生代真蕨植物系统研究的深度和水平。